In Runeroots TD, no two runs are the same! The enemy path, resources, perks, and even the enemies you are facing are randomized. Learn your enemies weaknesses, and react by building the perfect defensive setup, or use your rune powers if you are in a bind!

There are a variety of upgrades you can pick throughout each run. Perks modify a specific tower type, items enhance a single tower, and Specials which are global modifiers drastically changing the way each run is played. Do you make one super tower? Or spread the perks across many different towers?

Each tower has two rune traits. Pairing up towers within the same rune trait grant them powerful bonuses. With 8 different rune traits, there are plenty of options to mix and match. Plan ahead to build the ultimate combo!

Level up your town as you play! The town evolves over time and unlocks new features to use on your next run.